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scholarly article in The Coleopterists Bulletin, vol. 63 no. 4, December 2009
Giant Dung Beetles of the Genus Heliocopris (Scarabaeidae) S Pokorný , J ZÍdek , K Werner . Format 24 × 28 cm, hardcover. 136 pp., 271 figs (over 213 color photographs and 58 line drawings). ISBN: 978‐80‐902734‐0‐5. Taita Publishers. Hradec… is a species.